Familia
SATYRIDAE
Triphysa
albovenosa Ershov, 1885 (= phryne auct.)
Triphysa
albovenosa sacha Korshunov, 1996 (fig. 1,
2,
3,
4).
Translation
of the original description:
HOLOTYPE:
a male. FWl. 18 mm. UPS is blackish-gray with a steel flash, without any
pattern, only dots or ocelli of the underside appear through slightly. The UNS
is as in T.
albovenosa, but there are small dots or ocelli in the outer field, the
second one of which on the HW even has a white pupil. There are three dots and
one ocellus on the FW, the HW has five small black ocelli between the veins, the
third from above being a dot. In some paratypes the dots are only on the HW, in
two paratypes they are hardly noticeable at all.
ALLOTYPE:
a female. FWl.: 15 mm. The UPS is grayish-white, the underside pattern is
appearing through the wing. On the UNS there are very small black dots: four on
the FW and five on the HW. The zone between the outer field and the base is
mottled due to gray spots of different sizes. In other females these gray spots
fuse to each other, in one female the UNS is as in the males,the dots are large,
especially on the FW.
MATERIALS:
The holotype: ♂ 7.06.1985, Yakutsk, the botanical garden (Dubatolov).
The allotype: ♀ 9.06.1985, the same locality. Paratypes: ♀ and 2
♂♂ 13.06.1968, Yakutsk, Chuhur-Muran;
♂ and ♀ 10.06.1974, Khaptagai, the Tamma River valley (Ammosov);
♂ 1.06.1977, at the settlement Oktemtsy; 3 ♂♂ 1 ♀
28-31.05.1989, Yakutsk, the botanical garden (Dubatolov);10 ♂♂
10 ♀♀ 7-9.06.1985, the same locality; a male
1.06.1985, 5 km of
Yakutsk on the road to Magadan, a bogged rivulet valley (Dubatolov).
Thus,
the range of a new ssp. lies within Yakutia. It is noteworthy that the
Suntar-Khayata Mt. Range in E. Yakutia is inhabited by different butterflies,
close to the ssp. tscherskii
Crum-Grshimailo, 1899. It has a
somewhat darker ground color on the UNS, the light veins being more conspiculus
as compared with other ssp. This ssp. ranges widely in E. Siberia, it was also
found in Transuralia and West Chukotka.
Origin:
Dopolnenie 1, Novosibirsk, 1996, pp.
30-31.
The
ssp. albovenosa
occurs in Priamurye and Primorye. The ssp. sacha
was described from the Prilenskoe Plateau. In the E. Yakutia and Suntar-Khayata
Mt. Range we find other butterflies similar to the ssp. tscherskii
Grum-Grshimailo, 1899
having somewhat darker background of the UNS in comparison with other ssp., the
light veins are clearer visible. The ssp. tscherskii is widespread in the E. Siberia and is found in
Zauralye and in the the western Chukotka.
SYSTEMATIC
NOTES. There are two names older than albovenosa:
dohrni Zeller,
1850 (the description was made for the unique specimen caught
supposedly in S. Russia although a Siberian origin seems more probable; this
specimen was lost in the 19th century) and nervosa Motschulsky,
1866 (the description was made for Japan, where Triphysa
is not present!). The diagnoses by Motschulsky and Zeller dont coincide, and
the diagnosis of the latter is similar to glacialis.
Coenonympha
glycerion (Borkhausen, 1788) (= amyntas Poda, 1761, iphis [Denis et Schiffermüller], 1775)
The
nominotypical ssp. occurs as far east as W. Altai. In the upper reaches of the
Ob River and further in the mountains there occurs the ssp. iphicles Staudinger,
1892; these butterflies have more or less perceptible ochre rings on the UPH
and enlarged spots with ochre borders on the UNH (in glycerion
they are yellowish). The ssp. heroides
Christoph, 1893 from the Lena R. basin [Korshunov, Ammosov, 1978] has rather
larger spots. The ssp. beljaevi was described from S. Primorye.
Coenonympha
glycerion beljaevi Dubatolov, ssp. n. (fig. 1,
2,
3,
4).
Original
description:
Coenonympha
glycerion beljaevi Dubatolov,
ssp. n. (Figs 21-22)
MATERIAL.
Holotype: ♂, Southern Primorye, Spassk Distr., Novovladimirovka village,
27.VI 1978 (Barkalov). Paratypes: ♀, the same locality. 27.VI 1978
(Barkalov); 3 ♂♂, 3 ♀♀, Southern Primorye, Anuchino
Distr. 1.3-21 km NNW of Chernyshevka village, 12-17.VII 1993 (Dubalolov,
Zmishcnko): ♀. 2-3 km N of Chernyshevka village, 5.VII 1995 (Dubalolov,
Dudko).
DESCRIPTION,
MALE. FWl. 17-19 mm. The UPS is evenly dark brown with a narrow reddish margin,
usually without any visible-through dark spots along the HW margin. The wing
base has the same color as the main part of the wing. The underside is of the
same color, but slightly lighter. The FW bears a small apical ocellus and has a
reduced white postdiscal band between the veins M1 and M2 (most paratypes have
no such band). The HW has a row of very large ocelli with wide black and orange
rings, therefore the orange rims fuse and form an orange band. Proximally to the
row of ocelli there are remainders of the white postdiscal band, consisting of 2
spots: an irregular one between the veins M2 and CuA with proximal tooth on the
vein M3 and a small one placed between the veins Cu1 and A2. There is a
narrow silvery band along the HW edge.
FEMALE.
FWl. 16-19 mm, the UPS is brown with light-orange margin, a visible-through
apical ocellus on the FW and orange ocelli along the HW margin. The pattern on
the UNH is as in males, but the white nuclei within the black ocelli are very
wide, up to 1 mm in diameter. The UNF has an apical ocellus and 2-3 ocelli in
the spaces 2, 3 and 4 and a well-developed postdiscal band between the stem R
and the vein Cu1.
DISTRIBUTION.
Russia: Primorski Territory (Siniy Mt. Range).
REMARKS.
The new ssp. differs from the South Siberian C.
g. iphicles Staudinger, 1892 by an usual absence of a visible-through
ocellus on the male UPH (figs. 21-22), which is well visible in C.
g. ipflicles (figs 23-24): the black rings of the UNH ocelli in C.
g. beljaevi are wide, their width is considerably larger than the
distance between the rings, whereas the width of these rings in C. g. iphicles is considerably less than the distance between
them. The orange rims of ocelli of C.
g. iphicles are not fused but well separated from each other. The
nominotypical ssp. C. g. glycerion (Borkhausen,
1788) from Europe and West Siberia has very small ocelli on the UNH, their
diameter often being less than the distance between them (figs 25-26). In
appearance the new ssp. is most similar to the Yakutian C. g. hercules Christoph, 1893
(described from Witim and Wilui). The UNH ocelli of the latter ssp. are
also large, with large white centers, but the centers are smaller and the orange
rings around the ocelli are narrower (figs 27- 28) and are not fused into a band
(at least in the males), as in the new ssp. The FWl. of the Yakutian C. g. heroines is much smaller (14-15.5 mm).
Origin:
Far East. Entomol. 1997, No. 44, pp.
8-10, Figs 21-28.
Coenonympha
amaryllis (Stoll, 1782)
From
the Urals across the S. Siberia the ssp. amaryllis
is widespread. The taxon rinda
Ménétriés, 1859
was described on the basis of an an aberrant specimen from the Amur valley. This
fact has ben was forgotten. In Seitzs work the rinda
is depicted differently; its image in the table doesnt even resemble rinda.
The available specimens from Priamurye and the E. Siberia are similar to the
Manchuro-Korean ssp. accrescens
Staudinger, 1901 (with a complete set
of ocelli). This ssp. was captured in the basin of Ussuri, as well. As to the
butterflies like rinda,
they are to be found, for example, in the basin of Zeya, but they are not so
frequent and differ by a distinct degree of reduction of the ocelli and others
elements of the pattern. The largest and most peculiar specimens of amaryllis
are described by us as the ssp. borisovi.
ooenonympha amaryllis borisovi Korshunov et Ivonin,
1996 (fig. 1,
2,
3,
4,
5).
Translation
of the original description:
...The
largest and peculiar butterflies among amaryllis
have been found in Pribaikalye, which are decribed as C.
amaryllis borisovi Korshunov et Ivonin, ssp. n.
HOLOTYPE:
a male. FWl.: 20 mm. The UPS is ochre-yellow. There are two large and two small
black dots on the FW and five large dots and a dark marginal rim on the HW. On
the underside there are very large, contrasted ocelli with silvery pupils and a
relatively wide silvery streak along the margins. The fringe is light. On the
most part of the sp. the butterflies have the underside without any pattern,
that on the underside is more dull, the ocelli are smaller than in borisovi.
MATERIALS:
The holotype: ♂ 24.06.1988, Baikal, the v. Listvyanka (Ivonin).
Paratypes: 2 ♂♂ 30.06.1983, the same locality (Yu. Timoshenko),
3 ♂♂ -24.06.1988, the same locality
20.06.1991, the Verkhne-Angarskii mountain range, the
environs of Severobaikalsk (Ivonin). All the butterflies were collected on
steppefied mountain slopes. The ssp. is named by the surname of a Decemberist
Borisov, Petr Ivanovich (1800-1854), the author of first aquarel portraits of
Lepidoptera of Baikal.
Origin:
Dopolnenie 1, Novosibirsk, 1996, p.
31.
We
introduce here the following improvements into the description of borisovi.
The
FW are 19-21 mm long. Some specimens have clearly discernible black points on
the FW and the HW, the other have less visible or almost imperceptible dots. The
largest dots are about 1 mm in diameter at the apex and on the ventral part of
the HW, where 2-3 dots are usually visible. Other points are sometimes as small
as twice and are hidden among the distinct ochre flakes. The quite narrow dark
border of the UPS is more appreciable on the HW; it is present on the FW, too,
although there are specimens whose border is hardly noticeable. A paratype
specimen has clearly visible dark spots on the fringe of the FW. The UNF has
single or double ocelli at the apex and two ocelli below, the HW with six ocelli
surrounded by ochre rings; they form an arc along the outer border. Between this
arc and the edge of the wing a silver sinuous band is visible, and the inner
edge of the arc has a white band. The white mark is located opposite the central
cell and near the lowermost ocellus (it is not clearly perceptable in all
specimens). The ocelli on the FW are generally framed the same way, but they
have no ochre rings and are enclosed in light scales. The large cores of the
ocelli seem to be silvery, especially seen in the bright light, but they are
actually white. It is necessary to add to the paratypes a male from Pad Chernaja
near Baikal from 3.07.1959. As a rule all specimens have ochre UNF and ash-gray
UNH. Furthermore, the genital apparatus of borisovi is quite
distinct, sufficiently so at times to justify separating it from the other
suibsp.On the figure one can discern a narrow and oblong subuncus, as well as a
specific form and size of the valva and of the aedeagus. One notes a certain
resemblance to the chinese ssp. of amaryllis, accrescens Staudinger,
1901. But its habitat is remote and the image in Seitz (tabl. 48f) nor only
doesnt coincide with our ssp., but is even notably different.
Origin:
Dopolnenie 2, Novosibirsk, 1998, p.
22.
Hyponephele
cadusina (Staudinger, 1881) (= chamyla auct., kirgisa auct.)
Among
the specimens I caught in the Katun River valley the ssp. gurkini has been described
as Hyponephele cadusina gurkini
Korshunov, 1995.
Hyponephele
cadusina gurkini Korshunov, 1995 (fig. 1,
2,
3,
4).
Translation
of the original description:
...A
new ssp. Hyponephele
cadusina gurkini Korshunov, ssp. n. is described on materials from
Central Altai, differing from the nominotypical one by the following features:
on the UPF the yellow field is pale, the dot between the veins Cu1 and Cu2 is
larger, the dark margin on the UNH is wider, the UNS is more evenly colored, the
dots at the ventral angle are smaller.
HOLOTYPE:
a male. FWl.: 20 mm. The UPS is brown, the HW is evenly colored, the FW has a
wide ochre-yellow spot on the outer field. In the apical part of this spot there
is a black ocellus with a light pupil and as small as twice black dot below it,
between the veins Cu1 and Cu2. A black androconial brand is covered by brown
scales along its fore and hind margins. The UNF is ground-yellow with a
relatively wide dark marginal band and a light-rimmed apical ocellus. The UNH is
gray with brown marble which does not form transversal bands, as is observed in cadusina.
MATERIALS:
the holtype 1st July 1974, Central Altai, a steppefied bank slope
of the Katun River near its junction with the Chuya River, (Yu.P. Korshunov
leg.). Paratypes: 2 ♂♂, 1st July 1974, the same
locality.
The
ssp. is dedicated to the memory of an Altaian painter Gurkin Grigori Ivanovich
(1869-1937)
Origin:
Dnevnye babochki, Ekaterinburg, 1995,
p. 121.
Erebia ligea
(Linnaeus, 1758)
The
butterflies from the Polar and Subpolar Urals and the S. Yamals have blind,
frequently prolated ocelli on the FW and reducted androconial spots, they look a
bit like the Lapland ssp. dovrensis Strand, 1902.
In the rest of the Urals as far as Prisayanye inclusive occurs the ssp. kamensis
Krulikowsky, 1909. This butterfly has a relative reduction of the white band on
the UNS of the HW. From the Altai as far as the Pacific Ocean the ssp. umonia
Ménétriés, 1859
(= ajanensis
auct., koreana Matsumura,
1928) is found. This ssp. is remarkable for yellow-brown coloring of
the narrower bands of the FW as compared to kamensis (fig.
1,
2,
3,
4)..
This ssp. was described on the basis of five females and twelve males in
Shrenks collection from the Khadzhi Gulf (nowadays Soviet Harbour). The
butterflies from Sakhalin similar to eumonia were described as takanonis
f. sachalinensis Matsumura,
1919. In Kamchatka the butterflies
are known as kamtschadalus Goltz, 1932:
their band on the FW is devided by veins and consists of separate spots.
A complex Erebia euryale (Esper, 1805)
We
adduce here a supplementary information and descriptions especially concerning
the eastern taxa of butterflies of the Erebia euryale and E.
adyte groups. The given publication ascertains the former information
[Korshunov 1995, 1996, 1998, 2000] about euryale
in thje E. Europe and in the the Uralian Mountains.
Erebia
euryaloides Tengström, 1869, stat. n.
The
taxon was described as Erebia ligea euryaloides Tengström,
1889.
The
original description by Tengström: Minor, alis anticis supra ocellis
obsoletis l. inocellatis, subtus fascia rufa bsin bersus indeterminata
diffusaque, alis posticis infra obscure brunneis fascia pallidiori (l. albida
luteo irrorata in fem.) ocellisque evanescentibus Infra limites Fenniae
politicos. [Tengström in Warren 1936:71].
NOTE.
According to Warren [Warren, 1936:71-72], its principal characters are as
follows: The bands on the upperside of the forewings are broken up into spots,
and this feature is more marked on the hinwings, the spots there often being
reduced in size and some of them wanting. Two very small black dots at the apex
of the forewings are all that remain of the usual black spots; in some specimens
even these two are lost also, but more frequently the can be descerned. The
underside of the hindwinds is of the isarica
type of markings, the antemarginal band usually fairly distinct, strongly dusted
with silver but without spots; a distinct basal band is usually present.
Occasionally the silver streak along the inner edge of the antemarginal band is
very clear, but even in such specimens it is typically a euryale
streak and not a ligea
one Of the other ssp. or forms of euryale, euryaloides perhaps resembles
segregata the most; on the whole, however, the euryaloides markings are much
more stable, and the absense of thq black spots, other than the two apical ones,
together with the more pronounced marking of the underside of the hindwings
(also without spots), give true evryaloides a fairly distinctive appearance.
Unfortunately I have not seen a female.
About
the geographical distribution of butterflies B. Warren writes only that ssp. euryaloides
in the Uralian Mountains only varies aberrationally from the Finnish race.
But the material from the Urals as well as from the western territories was in
that time very scant, and Warren was caused to write that the limits of the
distribiution of euryaloides in Finland are very little known, but it is not
likely that it is very localized. It extends eastwards from Finland into Russia,
and I have seen specimens fros as far as north as the northern end of Lake
Onega. Far to the east the insect reappears un the Ural Moumtains, and of its
range there little or nothing is known This, however, is the most easterly
locality known for euryale.
The
result of investigation of the available materials of butterflies united by
Warren in the ssp. euryaloides
is as follows.
Firstly,
there are reasons to raise the status of euryaloides
up to a specifical one (see argumentation below). There are some good ssp.
inside of the sp. euryaloides.
Secondly,
the Uralian Erebia iremelica must be separated from euryaloides whose specific
status is based on the structure of male genitalia, on the antemarginal band
shape and on common fly of two sp. in some Uralian regions. The features assumed
by Warren as typical for euryaloides, characterize
postulated by us ssp. flaveoides,
zhuravskyi and taiga;
they, however, dont correlate with Erebia iremelica features.
Thirdly,
the nominotypical ssp. E.
euryaloides euryaloides doesnt occur in the NW taiga zone from Europe
as far as the Urals, the tundras beyond the the polar circle and in the Ural
Mts.― it follows from the original Tengströms description of
females with a golden-yellow UNH band. The question about the area of the
nominotypical form remains open: according to Tengström, it flies within
the state boundaries of Finland, but it is not clear which part of this country
within the XIX s. limits could be meant. By ty way E.
euryaloides is not indicated for Kola pen. The extreme trustworthy
western point of the caught of E.
euryaloides is Kosmozero, Medvezhjegorski reg., Karelian ASSR (the north
end of Lake Onega) [Warren, 1936, table, fig. ]. It is not excluded that Karelia is
the extreme eastern point of the
area of E. euryaloides. It is necessary to find out its exact area and
at least its presence within the contemporary Finland.
Fourthly,
the described by Poppius polar f.
arctica results an ecolofical form of E. euryaloides taiga (it is genetically analogous to polar forms
of E. e. flaveoides) ― it follows from the geographical
distribution of females with silver-white and yellow UNH band. The f.
arctica and similar tiny specimens of flaveoides differ from
their southern analogues in size only, being their tiny copies.
Translation
of the original descriptions:
Erebia
euryaloides zhuravskyi Nikolaev et Korshunov, ssp. n.
HOLOTYPE
― ♂. Body black, antennae and legs dark-brown from above, whitish
from below. The UPS brown. FWl 21 mm. The band is rusty-red, divided by thin
dark-brown veins; it is placed between R5―2V (almost reaching 2V). It is
evenly narrowing from the inner side downwards to the ventral edge forming a
blunt peg betwen Cu2―2V. The band at outer edge is flat. There are
blind ocelli betwen M1―M2 and M2―M3. The UPH band is colored
identically, it is between Rs―M3. 2 rusty-red spots (colored darker that
the band) are between M3―Cu1 and Cu1―Cu2. The first one is narrower
than the band, the second one is narrower than the first. Ocelli (dots) are
absent.
The
UNS are dark-brown. A transversal rusty-red band passes throug from Rs as far as
2V, it is diffused in its inner edge between M3―Cu2. The lower part of the
central cell and the discal zone between M3―Cu2 are rusty-red (and are
darker than the band). The band bears black dots betwen M1―M2 and
M2―M3. The background is a little darker than this of the UNF. The band is
silver-gray between the costal edge and the vein 3V, its outer edge is diffused,
the inner one is marked with silver-gray scales. A small silver-white triangular
spot (streak) is between the band and the central cell over M3; a similar
darker and smaller spot is on Cu2. Along the outer edge of the band between
M2―M3 and Cu1―Cu2 are two red spots with distinct dark dots in the
middle. The fringe is chequered, with alternating of equal white and brown
parcels.
The
valva tip is stretched, bearing short teeth at the front and from above. The
apical edge is gentle-sloped bearing rather evenly posited thick teeth of the
average size. The dorsal edge bears sparse large teeth whose dots form a
straight line. The valva hair is thin, mostly short, the brush on the top
of valvae isnt dense.
ALLOTYPE
― ♀. The background of the UPS is dark-brown. FWl 21 mm. The shape
and coloring of the outer zone band is like in the male. The seeing ocelli
are between M1―M2, M2―M3, a blind ocellus between Cu1―Cu2. 5
separated postdiscal rusty-red spots between R2―Cu2 are on the UPH, the 5th
spot includes a seeing ocellus. The 5th spot is round, the
other are roundish. The 4th and 5th spots have the same
width, they are narrower than the 3th one.
The
UNF background is dark-rusty-red. The costal and outer borders are dusted with
sparse silver-white scales. The band (R2―2V) is rusty-red (it is lighter
than the background), very light between R5―M3; its inner edge is diffused
between M3―2V. The UNH background is brown. The basal band is
silver-white, consisting of sparse scales. The passing through the whole wing
posdiscal band is silver-white, its inner edge is distinct, bearing a
protuberance over M3; the later almost reaches the central cell. The outer edge
of the band is suffudes. 3 rusty-red spots are along the outer edge between
M1―M2, M2―M3 and Cu1―Cu2. The seeing ocelli are between
M2―M3 and Cu1―Cu2. The fringe is like in male.
MATERIALS.
Holotype ― ♂, the Urals, Kosvinski Kamen mt., the top of
podgoltsovy zone, 15.07.1974 (Yu. Baranchikov). Allotype ―
♀, the Urals, env. of Kytlym, spruce-fir-cedar forest, 18.07.1974 (Yu.
Baranchikov). Paratypes ― 4♂, 10.07.71, Katav-Ivanovsk (Sablin);
1♂, 12.07.65, 1♂, 17.07.65, ibidem (Sablin); 1♂ 15.07.74,
2♂ 17.07.74, Kosvinski Kamen mt., the top of the predgoltsovy
zona (Yu. N. Baranchikov).
DISTRIBUTION.
Mountains of the North, Middle and South Urals (to the N as far as 62 N).
ETYMOLOGY.
The ssp. is named in honour of the researcher of the Northern Preduralye Andrei
Vladimirovich Zhuravsky (1882―1914).
Erebia
euryaloides taiga Nikolaev et Korshunov, ssp. n.
HOLOTYPE
a male. Body black, antennae and legs dark-brown from above, yellowish from
below. UPS brown. FWl 21 mm. The band is rusty-red, divided by dark-brown veins,
it is placed between the veins R5 and 2V. The band between M1―M2,
M2―M3 is a little wider than between the other veins. The large blind
ocelli are between M1―M2 and M2―M3, there is a hardly seen black dot
at the outer edge of the band between Cu1―Cu2 on the right wing. The outer
edge of the band is flat. The antemarginal band on the UPH consists of 5
separate rosty-red spots betwen Rs―Cu2, therefrom the spots between
M1―M2, M2―M3 are more stretched. Ocelli (dots) absent.
The
UNS is dark-brown. There is a rusty-red band on the UNF between the veins
Rs―2V, its inner edge is diffused between Cu2―2V. The lower part
of the central cell and the discal zone between V2 and the inner edge of the
wing are rusty-red (a little less brigher than the band). A seeing ocellus
is between M―M2, a blind one between M2―M3 (the later is smaller
than the first). The background of the UNH is equal to UNFs one. The
antemarginal band is absent. The outer edge is a little lighter than the basal
one due to the dusting of sparse goldish scales. Silver-white angular spots are
on the inner edge of the outer zone, their tips are directed to the base
of the wing, they are situated on the veins Rs, M1, M2, Cu1 and Cu2. A
silver-white streak is above the vein M3, its tip reaches the central
cell.
The
fringe of wings is motley, with an alternation of equal in size white and brown
parcels.
The
construction of valvae is identical to E.
e. zhuravskyis one.
ALLOTYPE
a female. Body, legs and
antennae are like in ♂. The UPS is dark-brown. FWl 21 mm. The UPF band is
similar to males one, the seeing ocelli are between M1―M2,
M2―M3. There is a supplementary little rusty-red spot between 2V―3V
on the UPH.
The
UNS underground is dark-brown. The rusty-red band is on the UNF and similarly
colored but a less bright zone like in ♂. The seeing ocelli
are between M1―M2, M2―M3, a blind one between Cu1-Cu2. The UNH outer
border is relatively lighter and whiter than the background. The
antemarginal band is silver-white, it has a distinct inner edge; its
protuberance is directed to the base of the wing, its tip almost reaches the
central cell; the protuberance is placed above the vein M3. The outer edge of
the band is diffused. 5 round rusty-red spots are stretched along the outer edge
of the band between M1―M3 and Cu1―Cu2. In the middle of them are
seeing ocelli. The rusty-red spots between Rs―M1, M1―M2 and
M3―Cu1 are small, irregular, without ocelli. There is a dim whitish basal
band. The fringe is like in the male.
MATERIALS.
Holotype ― ♂, Komi Rep., Ust-Vymski reg., Mikun mt., a medow,
plain part of the middle taiga, ~ 62 N, 25-27.07.1999. Alloptype ―
♀, ibidem, 25-27.07.1999. Paratype ― 1♂, 3♀ ―
ibidem, 25-27.07.1999; 3♂, 1♀ ― W. Pritimanye,
Arkhangeksk distr., Leshukonskoe v., ~ 64 N., VII.1980 (A. Bobretsov).
DISTRIBUTION:
taiga in the NE Europe (and tundra ― the ecological f. arctica Poppius, 1906).
BIOLOGY:
According to A. G. Tatarinov, it is a sp. with a biennial generation. The
imagines appear in the environs of Ukhta (middle taiga) only in the even years.
Nevertheless, K. F. Sedykh affirms that the butterfies still appear here in the
odd years, but their number is scant and they fly earlier. The period of flight
is late June ― early August. The butterflies feed nectar of buttercup,
valerian, .,
thistle, .,
..
Food-plants are various wild cereals. [Tatarinov, Dolgin, 1999:110].
ETYMOLOGY:
the ssp. is named in according to the place of the inhabit.
Erebia
euryaloides flaveoides Korshunov et Tatarinov, 1996, stat. n. (fig. 1,
2, 3,
4,
5, 6,
7,
8,
9,
10,
11, 12).
The
original description of the ssp. [Korshunov, 1996, 1998]; the supplementary
features are given between the square brackets.
flaveoides
Korshunov et Tatarinov, ssp. n.
HOLOTYPE
― a female. FWl 21 mm. There are two small brown spots on the
brown-yellowish band on the UNH: the upper one is opposite the hollow in the
band before the top of the central cell, the lower one is between the veins Cu.
[The paratypes may have one of these spots or nothing but the tracks (shades)].
ALLOPTYPE
― a male. FWl 20 mm. The UNH band is reduced, a white small spot is seen
where the female has a hollow, and sparse white scales above and below it. [The
UPF has a dark rusty-red band between R5―2V, its inner border is evenly
narrowing downwards. The outer band edge is flat. The UPH band is placed between
M1―M3, two roundish spots are between M3―Cu1 and Cu1―Cu2, a
hardly seen spot of the same color is between Cu2―2V. 2 blind dots
(ocelli) are only between M1―M2 and M2―M3. The valvae have a
shortened tip. The teeth tips on the dorsal side form a straight line, like in
ssp. zhuravskyi and taiga].]
[DISTRIBUTION.
The Polar, Subpolar and North Urals (as far as 62 N)].
[BIOLOGY.
According to A. G. Tatarinov, the imagines occur every year in the Urals
[Tatarinov, Dolgin, 1999:110].
MATERIALS.
Holotype ― ♀, 15.07.1995, Subpolar Ural, midstream of M. Patok rr.
Allotype ― ♀, 16.07, ibidem. Paratypes ― 9 ♀♀,
15-17.07, ibid.; [3♂, nat. p. Yugad-Va, Kozhim r., ~ 66 N, grassy
larch forests, 10-12.07.2000].
NOTE.
E.
e. taiga and E. e. zhuravskyi differ groundly in the structure of the UPF
band: the band of the latter sp. has its own distinct narrowing downwards to the
ventral edge of wings; the same narrowing in E. taiga is less noticeable.. E.
e. flaveoides has a slightly smaller dimensions; the coloring of the
female antemarginal band is yellow in flaveoides,
silver-white (silver-green) in zhuravskyi
and taiga.
In
according to A. G. Tatarinov small butterflies occur in the Polar Urals and
in the eastern part of the Bolshezemelskaya Tundra; ocelli are absent
frequently on their wings, the postdiscal band on the female UNH can be yellow
as well as white. The taxonomical situation of the form arctica
Pop. remains unclear; it was described from the Kanin Pen. and is
characterized by small size, reduced ocelli and a white band in females
The butterflies from the Transpolar Ural and the eastern part of the
Bolshezemelskaya Tundra are closest to this form. It is not excluded that
they must be considered as a separate ssp. [Tatarinov, Dolgin, 1990:110]. But
according to the geographical distribution of the dwarf form whose females have
a silver-white or yellow UNH band, we incline to assume them as transpolar
ecological forms of the ssp. E. euryaloides taiga and E.
euryaloides flaveoides, respectively. The analogous ecological dwarf
forms with similar features are found in other butterflies of the euryale-group,
e. g. in the forms like Erebia
euryale syrmia f. nana from
the High Carpathians [Warren, 1936].
Our
materials present only tiny copies of E.
euryaloides flaveoides. FWl is
17-18 mm. The learned Ural specimens, in accordance to Poppius original
description of arctica
from Kanin, the rusty-red spots are more or less reduced and have not ocelli.
The UNH white pattern along thre outer edge of the row of small rusty-red spots
is weakly distinct in males, only sometimes occur butterflies with a
well-developed white band (ab. boreomontanum
Sedykh, 1977). The valva tip has an
average length, the dorsal part is like in flaveoides,
taiga and zhuravskyi.
Two front spots on the UNF in ♀♀ bear white ocelli surrounded with
black rings. A wide silver-white (or ochre-yellow) band is well distinct in the
outer zone of the UNH, its inner edge is limited by a distinct roundish-broken
line of the adjacent dark stripe which is covered with white scales in the basal
zone of the wing. The females with a yellow band (parallel with white band
females) are noticed only in the mountains (according to Tatarinov, only
white-band ♀♀ inhabit the Bolshezemelskaya Tundra).
MATERIALS.
2 ♂♂ ― 23.07.1960, forest-tundra, near Labytnangi t., 1♂
1♀ ― 07.1963, Polarny Ural st., railroad
Vorkuta―Labytnangi, 1♂ ― 17.06.1966, Labytnangi (K. Sedykh), 2
♂♂ ― 16, 26.07.1972, 107th -109th km of
Seida―Labytnangi railr. (V. Makhat), 2♂♂ ― 18 and
27.07.1972, Polar Ural, M. Paypudin r. valley (V. Makhat).
Erebia
iremelica Korshunov, 1995, stat. n.
HOLOTYPE
― ♂. Antennae and legs are dark-brown from above, gray-whitish from
below, body black. Fringe gray-whitish with interspersed with black and white of
the background.
FWl
20 mm. The UPS are dark-brown with rusty-red postdiscal rows of spots between
veins. The UPH 5 spots band betwen R5―Cu2, the inner sides of the spots
1―3 are stretched towards the base, the 1st spot is from the
outside shorter then the 2nd and 3rd ones. The 4th
spot between M3―Cu1 is noticeable shorter and narrower than the 5th.
The black dots are in the dots 2, 3 and 5. The UPH band consists of 5 spots and
is stretched between Rs―Cu2; the sots 1 and 4 are noticeably shorter than
the rest, there is a the black dot only in the 5th spot. The UNS
background is dark red-brown. The UNF band is continuous, narrowed from the
upper side between the veins R5―M1 and A―Cu2 and from the basal side
between M3 and Cu3. The inner side of the band is diffused between M3―Cu2.
The lower half of the central cell and a part of the wing from the cell as far
as the band (between M3 and Cu2) is rusty-colored. There are black dots on the
band between M1―M2, M2―M3, Cu1―Cu2 and separate small
rusty-red spots between M1―Cu2 in the outer zone of the UNH. The spots
between M1―M2 and Cu1―Cu2 are roundish and bear seeing ocelli.
There are ochre-colored slightly rimmed by rusty-red streaksbetween the veins
M1―M2 and M3―Cu1. A short silver-white streak is over the vein M3
between a harly seen silver-white band and the central cell.
The
valva top is narrow, stretched, having a mace-shaped thickening on the top,
covered with small teeth. The apical edge is sloped, covered whith chaoticly
posited average teeth. The dorsal edge is bow-shaped, it is narrowing towards
the back part; the dorsal teeth are large, their length is various. The valvae
are covered with thick hair of average length, the longest hair is in the dorsal
part and on the top where a hair-brush is formed.
The
male variability among the paratypes concerns the following features: a)
presence/absence of ocelli on the UPS. A male has UPH ocelli in the spots
3―5, they are seeing in the spots 3―5. Another specimen has a
seeiing ocellus in the 2nd spot on the UPF, while a hardly seen
blind one is in the 3rd spot on UPH; 2 specimens have ocelli only in
the 2nd and 3rd spots on the FW, etc.; b) the brightness
of the silver-white band on th UNH. Some butterflies have bands more distinct
than the holotype, another have a hardly seen band; c) shape of the dorsal edge
of valvae. The dorsal edge always has a typical feature (differing iremelica
from euryaloides) ― a bow-shaped protubrance, similar to E.
jeniseiensiss one (differing from the later by chaotic disposition of
teeth and diversity of teeth shape). Many specimens have a regular
bow-shaped swelling, some butterflies have several protuberances on the curved
dorsal edge.
ALLOTYPE
― ♀. UPS dark-brown. The band consists of separate rusty-red spots
separated by veins. FWl 22 mm. The UPF band is between R2―Cu2. The 2nd,
3rd and 5th spots contain ocelli (a blind ocellus between
Cu1―Cu2). The 1st and 4th spots are narrower and
shorter than the rest. The UPH band has 4 spots without ocelli between
M1―Cu2. The 1st and 3rd spots are narrower and
shorter as the 2nd and 4th. The inner edges of the 1st
and 2nd spots are slightly stretched towards the base.
The
UNF is rusty-red, the costal border up to the vein K5 is silver-grayish (exscept
its submarginal part), the coloring is the most intensive above the band. tHE
The submarginal zone is thickly dusted with silvery scales between the costal
edge and M3. The band is rusty-red, lighter than the background, noticeably
narrowed between M3―Cu1. It bears 3 ocelli (between M1―M2,
M2―M3 and Cu1―Cu2), the Cu1―Cu2 ocellus is blind. The UNH
background is red-brownish, dusted with silver-white scales. The outer zone band
is silver-white, a peg-shaped projection lies over M3, its tips almost touches
the central cell. There are tracks of red-brownish spots between M2―M3 and
Cu1―Cu2 and a a silver-white basal band.
The
wing fringe is white with dark-brown uncrements.
The
females with silver-white or yellowish antemarginal UNH band are fixed between
the paratypes.
A
male have UPH ocelli in the spots 3―5, the ocelli in the 3th and 5th
spots are seeing. Another male has a seeing ocellus on the 2nd
spot on the UPF, while the UPH have hardly seen traces of blind ocelli in the 3rd
spot; two ♂♂ have ocelli only in the 2nd and 3rd
spots on the UPF, etc. (it is an important distinctive feature of E. iremelica, because as a
rule all the ssp. of E.
euryaloides have only two blind ocelli between the veins M1―M3 on
the UPF in males).
MATERIALS.
Holotype ― ♂, 26.06.95, Tschelyabinsk distr., Kusikhinsky reg., nat.
park Zyrat-Kul (V. K. Zinchenko). Allotype ― ♀, 26.06.95,
ibid.. Paratype: 9♂, 26.06.95, ibid.; 2♂, Katav-Ivanovsk, 16.07.67
(Sablin); 1♂, 14.07.63, 2♂, 12.07.65, 1♂, 3.07.65, 1♀,
12.07.65, ibidem (Sablin); 1 ♀ ― Chelyabinsk disr., Iremel,
meadows on the way towards Tyulyuk, Karachayka r., 15.07.1966 (E. Yu.
Zakharova); 1♀, 5♂ ―Chelyabinsk distr., Iremel, 13.07.1996;
7♂ ― Sverdlovsk distr., Visimski st. preserve, 20-22.07.1999
(V.M.Yudin); 2♂ ― the Urals, env. of Kytlym, spruce-fir-cedar
forest, 18.07.74; 1♂, Ural, Kosvinski Kamen mt., mountain forest
zone, 11.07.74 (Yu. N. Baranchikov).
NOTE.
According to Yu. N. Baranchikov [Baranchikov, 1980: 218], the butterflies were
gathered in the summer 1974 from 6 until 25 July mainly on the slopes of the
Kosvinski Kamen Mt. and in taiga in the environs of Kytlym. Thus, both
sp. were caught in (E.
iremelica
E. euryaloides
zhuravskyi)common in the env. of
Katav-Ivanovsk (S. Ural) and Kytlym (boundary between the M. and N. Urals). A
typical E
iremelica (labelled Ural) is shown by B. Warren [Warren, 1936, pl.
62, fig. 629, 635]; the genitalia of this male see ibid., pl. 25, fig. 249.
DISTRIBUTION.
Mountains of the M. and S. Urals.
Some
peculiar male specimens were found out among numerous buttreflies of euryale-group caught by A.
S. Nikolaev in July 1995 in SE part of the Ukranian Carpathians, along the NE
slop of Svidovec Mt. Range, a southern spur of the Gorgan Mts., in the
polinonas Dragobrat (NE slope of the Blyznyca Mt.) and Shasa (between the
Ungareasca and Tîmpa Mts.) and along the road between them, on the height
1300―1600 m. The main part of butterflies from the Svidovec Range. belong
to Erebia
euryale Esp., asssumingly to its nominotypical ssp. (although there are
some grounds to attribute them to a separate E. Carpathian ssp.). The mentioned
peculiar males have distinctive outer (pattern) and anatomical (valva
shape) features, which show their special status, in our opinion. Outer features
of a caught female can be proposed as an allotype to the assumed new taxon. The
distinctive features and the common fly togeter with E. euryale make possible
the giving them a status of a bona sp.
Erebia polonina
Nikolaev, sp. n.
HOLOTYPE.
A male. FWl 21 mm. Body black from above, dark-brown
from below, legs gray-brownish. Antennae dark-brown from above,
dirty-white from below. The wing fringe has alternating dark-brown and white
spots, dark-brown under Cu2 on the FW and 2V on the HW with white spots opposite
to red spots of the band. The UPS are dark-brown. The postdiscal band consists
of separate rusty-red spots, 7 spots between the veins R5―2V on the UPF, 5
spots between Rs―Cu2 on the UPH. The spots between M1―M2,
M2―M3 and Cu1―Cu2 on the UPF are oblong with blind spots (dots). Two
vertically posited dots are betwen Cu2―2V. The spot between M3―Cu1
is shorter than the previous and the following, it has th shape of a square put
on the corner. A narrow spot without an acellus is between R5―M1.
Hindwings: the first 3 spots are oblong, without ocelli. The spot betwen
M3―Cu1 is narrow, of an irregular oval-rombical shaped. The spot between
Cu1―Cu2 is roundish, with a cblind ocellus. The background of UNF is
dark-brown, the lower half of the central cell and the space between
M3―Cu2 from the postdiscal band as far as the base are rusty-red. The band
is continuous, rusty-red, lighter at the costal edge, narrowed from within and
from the outside between M3―Cu1 and Cu2―2V. The background of the
UNH is red-brownish. The postdiscal band is continuous, gray-brownish, dull,
dusted with sparse goldish scales. There is a white triangular streak on the
innerside of the band above Cu2, whose tip reaches a half way from the band as
far as the central cell. The basal band is weakly distinct (descernible during
turning the butterfly relative to the souce of light). The blind ocelli are on
the band between M1―M2, M2―M3, Cu2―Cu3 (FW) and between
M2―M3 and Cu1―Cu2 (HW). The dorsal edge of valvae is bow-shaped, it
is covered with irregularly posited teeth of an average size, the tips of teeth
dont form a straight line.
?
ALLOTYPE. A female. FWl 21 mm. Body, antennae, legs and fringe like in the
♂. The UPS is dark-brown. A continuous rusty-red band is between
R5―Cu2, it is cut through by thin veins, two vertically posited dark-red
spots are between Cu2―2V. The ocelli are inside of red spots between
M1―M2, M2―M3 (seeing), Cu1―Cu2 (blind). The UPH band
consists of separate dark-red spots between Rs―Cu2. The spot between
M3―Cu1 has an irregular shape, without an ocellus; the spot between
Cu1―Cu2 is roundish, a little larger than the previous, bearing a blind
ocellus. The UNF is like males one; the light rusty-red postdiscal band is
narrowed from within between M3―Cu1. A whitish dusting is seen over the
band on the costal border. The inner border is gray-brownish. The UNH is
red-brownish, dusted with sparse goldish scales. The postdiscal band consists
from sparse silver-withe scales on the outer edge, the inner edge is bright
silver-white, it forms peg-shaped protuberances, their tips are directed towards
the base. The ocelli on the band are betwen M1―M2, M2―M3
(seeing), Cu1―Cu2 (blind) (FW); between M2―M3, Cu1―Cu2
(seeing), M3―Cu1 (blind). The basal band is weakly developed,
silver-white along the outer edge.
MATERIALS.
Holotype ― ♂, Ukr. Carpathians, Svidovec mt. rng., pol. Dragobrat,
1300 m, 20.07.95 (A. S. Nikolaev). ? Allotype ― ♀, ibid., pol.
Shasa, 22.07.95 (idem). Paratypes ― 5♂, Ukr. Carpathians, Svidovec
rng., pol. Dragobrat, 1300 m, 20.07.95 (A. S. Nikolaev); 5♂♂, ibid.,
the road between pol. Dragobrat and Shasa, 1600 m., 21.07.95 (idem);
4♂♂, ibid., pol. Shasa, 1400 m, 22.07.95 (idem); 1♂, Gorgany
mts., v. Bystrica, a tall-grassed water-meadow, 1000 m, 11.07.67 (I.
Kozakevich).
ETYMOLOGY:
The sp. is named aftre the cardinal dwelling place, the E. Carpathian gently
sloped mountains (poloninas).
SYSTEMATICAL
NOTES. The main differences of E.
polonina from the E. Carpathian euryale
consist in the shape of spots forming the postdiscal band (see the description
of the holotype): there is no spot reduction in euryale (resp. a sizable
narrowing of bands) between M3―Cu1; if it occurs in aberrations, the spot
between Cu1―Cu2 is amaller than the spot between M3―Cu1, or the
firts is equal to the second. Apart of it, the inner edge of spots between
M3―Cu1 in euryale
with a normal band (not reduced and not diffused, as in some aberrative
specimens) has a form of a straight line or a hollow (this edge is curved
outwards, roundish or triangular). The dorsal edge of valvae of euryale
is covered with large, average and small teeth, besides the tips of large teeth
form a straight line, while polonina always has a dow-shaped
protuberance, this bow can be divided in two by a little hollow.
Formerly
polonina
was known only from Svidovec (a southern spur of Gorgan). Later in the
Zoological museum of IESA SD RAS a similar male was found; it was caught in
Bystrica (Nadvornyanski reg., Ivano-Frankovsk distr., leg. I. Kozakevich) placed
right northwards from Svidovec on the NE slope of Gorgan. Among our materials
from another Carpathan localities (N slopes: Turovka, Rozhnyatovski reg.,
Iv.-Frank. distr., Semakovo/Konyatin, Putilski reg., Chernovcy distr.; S
slopes: Lugi, Kozmeshchek, Rakhovski reg., Sinevir/Sinevirskaya Polyana,
Mezhgorski reg, Zakarpatskaya distr.) Erebia
euryale is the unique representant of the group in question.
It
is noticeable the indication to E. euryale adyte supposedly inhabiting the E. Carpathian massive
Czarnohora, the SE spur of the Gorgan Mountains [Krzywicki, 1966:19]. It seems
that some specimens of E.
polonina were interpreted as adyte.
The
butterflies of the euryale-group were hitherto considering as pertaining to an
indivisible sp. Erebia
euryale (Esper, 1805), whose
several taxons were were joined with Erebia
ligea (Linnaeus, 1758). These facts are cited in B. Warrens monograph
[Warren, 1936], where all the taxa were presented as ssp. (4), forms (20) and
aberrations (12) namely of E. euryale. This system was defined more exactly
during the time, in particular Warrens forms obtained the status of ssp.,
especially for the Alpine [Roos & Arnscheid, 1976; Arnscheid & Roos,
1977] and Uralian areas [Korshunov 2000]. The information about the eastern part
of euryales area, wich was presented weaker by Warren due to
the absence of relevant materials, can be introduced only now as a result of
thorough investigation of specimens after their intensive capture by S. L. and
A. S. Nikolaevs in the Ukranian Carpathians, by M. P. Sablin, A. G. Tatarinov
and the other colleagues in the Uralian region.
The
analysis of wing patterns, genital apparati, localities and way of life of imago
permits to make conclusions as follows.
The
taxa is divided in two complexes ― euryale proper and adyte.
Now
we present this information as a whole.
The
contemporaneous classificatgion of the ssp. of E.
euryale can be presented as it follows (in the given list the
infraspecific Warrens forms having separate areas, are treated as ssp.;
Warrens ssp. acquire a range of infraspecifical groups). This list formally
contains ssp. not cited by Warren (independently from their status in relevant
publications).
Formal
similarities exist among Erebiae of the groups 1―3, on the one hand (euryale-complex),
and Erebiae of the groups 4―6, on the other hand (adyte-complex).
Namely, the Erebiae of the adyte-complex
have a narrowed outer edge of the band on the FW resp. reduced spots between
M3―Cu1, on the one hand, and a bow-shaped dorsum of valvae, on the other
hand. The butterflies of the euryale-complex
can have sporadically the mentioned first feature (in the groups ocellaris,
segregata,
antevortes,
in aberrative forms of flaveoides,
isarica syrmia), but the
second (more important) feature is permanently absent: all known to us males of
the euryale-complex
have only the straight dorsum (see tabl.. , fig
). The fact is important that the butterflies of the groups 1―3
(euryale)
and 4―6 (adyte) can coexist (namely adyte and isarica,
adyte
and segregata;
euryale
and polonina;
iremelica
and zhuravskyi),
whereas among the same complex there are no geographical meetings or
common fly of butterflies (the coexistence of euryale, isarica and syrmia
in the E. Carpathians, assumed by Warren [Warren, 1936], has no factual
confirmation).
The
status of a sp. (opposite to ocellaris) was already proposed for adyte [Roos &
Arnscheid, 1976; Arnscheid & Roos, 1977] ― to be certain, it was said
that these taxa are semi-sp. in Lorkovics terms. Indeed the
semi-sp., i. e. close sp. distributed in adjacent, almost or fully
separate areas (similar to the sp. Erebia tyndarus, E.
nivalis and E. calcaria within the tyndarus-group),
are not transitional taxa between sp. and ssp., but the true sp. All the
more, the shown above intersection of areas of the ssp. of euryale
indicates their specific status. Adyte
is also opposed to isarica,
if we dont consider segregata
as a transitional form (Mischform) from adyte to isarica,
like it is done in [Arnscheid & Roos, 1977], but as a peculiar ssp.
outwardly similar to ocellaris;
its shape of valvae is not known to us.
The
classification of taxa among the complexes is more complicated. Tramelana, isarica,
euryale, syrmia
ocellaris have
adjacent (not intersected) areas within the euryale-complex (the
groups 1―3). Antevortes has a separate, but not very remote area. The taxa of
the 4th group inhabit a quite separate area (NE Europe, The Urals)
and have a number of specific common features. There are two possibilities: to
assume all the taxa of the fourth group as ssp. of euryale,
or as ssp. of a separate sp. euryaloides,
which forms together with euryale
a natural complex (group of sp.); the same refers to antevortes: Erebia
(euryale)
euryale
Esp. (with ssp. euryale, isarica,
tramelana,
syrmia;
segregata,
ocellaris),
Erebia (euryale) antevortes
Vrty. (with ssp. antevortes, cantabricola, pyraenaeicola)
and Erebia
(euryale)
euryaloides
Tgstr. (with ssp. euryaloides,
taiga,
flaveoides,
zhuravskyi).
The second desision is supported by the fact of presence in butterflies of euryaloides-group
of common features not peculiar to another euryales
(in particular, the preservation of only 2 blind ocelli on FW of males). The
same reason permits us to suggest the separate status of E.
(euryale) antevortes.
The
same scheme works in the case of the adyte-complex
(the groups 4―6): Erebia
(adyte)
adyte
Hbn. ( etobyma
brutiorum);
Erebia (adyte)
polonina Nikolaev;
Erebia (adyte)
iremelica
Korshunov.
We
show this information as a whole:
I.
EURYALE―COMPLEX
1)
euryale-group
ssp.
tramelana
Reverdin, 1918 ― Jura
ssp.
isarica
Heyne, 1895 ― Savoy, Bernese Alps, the northern part of the C. Alps, E.
Alps, Julian Alps, Karawanken
ssp.
tatrica
Strand, 1915 ― W. Carpathians
ssp.
euryale
(Esper, 1805) ― Sudets, Bieskids, NW Eastern Carpathians
ssp.
syrmia
Fruhstorfer, 1909 ― Balkans, Transilvania,
S. Carpathians, southern part of the E. Carpathians
ssp.
ocellaris
Staudinger, 1861 ― southern part of the E. Alps (the centre of the area
― Dolomites)
ssp.
segregata
Reverdin, 1918 ― Gurnigel, Moleson, Vaud, Berner Oberland
2)
antevortes-group
ssp.
antevortes
Verity, 1927 ― High Pyrenees
ssp.
cantabricola
Verity, 1927 ― Cantabrian Mountains
ssp.
pyraenaeicola
Goltz, 1930 ― E. Pyrenees
3)
euryaloides-group
ssp.
euryaloides
Tengström, 1869 ― Finland, Karelia (?)
ssp.
flaveoides
Korshunov et Tatarinov, 1996 ― Polar, Subpolar and Northeern Urals (as far
as 62 N southwards)
ssp.
taiga
Nikolaev et Korshunov, ssp. n. ― taiga of the NE Europe;
Malozemelskaya, Bolshezemelskaya tundra, Kanin pen. ― the
ecological f. arctica Poppius, 1906
ssp.
zhuravskyi
Nikolaev et Korshunov, ssp. n. ― Middle and Southern Ural
(as far as 62 N northwards).
II.
ADYTE―COMPLEX
4)
adyte-group
ssp.
adyte
(Hübner, 1818-1822) ― the southern part of the W. and C. Alps
ssp.
etobyma
Fruhstorfer, 1909 (? = phoreta
Fruhstorfer, 1918) ― Maritime Alps
ssp.
brutiorum
Turati, 1911 ― Apennines
5)
polonina-group
ssp.
polonina
Nikolaev, sp. n. ― E. Carpathians (Gorgan Range only?)
6)
iremelica-group
ssp.
iremelica
Korshunov, 1995 ― M. and S. Urals.
It
is noteworthy, that the complexes euryale and adyte
differ not only by morphological but also by biological characteristics. The sp.
of the complex adyte
are butterflies of high-mountains, they fly on the heights 1000―1700 m in
all habitats (they sink or climb very rarely and apparently dont have
dwarf forms); they prefer the lower borders of conifer-forests. In
contrast the butterflies of the complex euryale
tyend to a relative eurybiosity in limits of mountain-taiga zone. For instance,
they occur in the Carpathians in the heights from 500 as far as 2000m, inhabit
also open grassy meadows; in the NE Europe they inhabit the mountain slopes as
well as the taiga an tundra. The transpolar and alpine euryales
occur in dwarf form. Meeting with E. ligea, E.
euryale can form hybrides with the former in the Carpathias (Bowtsarya
Mt. Range, ca. 650 m., S. Nikolaevs collection): a male has a typical ligeass
valva and euryales the right one (see the drawing). Analogous
facts noticed in the Urals A. G.
Tatarinov (personal communication).
Erebia ajanensis
Ménétriés, 1857 (fig.
1, 2)
The
taxon ajanensis
was described on the basis of a female. The ssp. arsenjevi Kurentzov
replaces the ssp. ajanensis,
1950 in the Sikhote-Alin Mt. Range,
whose butterflies usually have an incomplete albescent band and all features of
the group vilderi.
The
butterflies from the northern Priokhotye are a little different. They were
described as Erebia
kosterini sp. n. It was later discovered that kosterini is a ssp. whose
imago tends to have somewhat oblong ocelli on the UNH together with a clear
albescent band through the entire wing. They also differ in the details of the
genital apparatus, as is evident in the drawings
The
translation of the original description:
Erebia
kosterini P.Gorbunov, Korshunov, Dubatolov, sp. n.
TYPE
LOCALITY: The Khindzha River on the Koni Peninsula (the Magadan Region)
BIOTOPES:
valley meadow plots between the thickets of the bush alder and the dwarf pine.
FLIGHT
PERIOD OF IMAGINES: observed in the second half of July.
HOLOTYPE:
a male: FWl.: 21.5 mm. The UPS is brown with a wide (3-4 mm) ochre-fulvous band
in the postdiscal area, which contains: a black dot at the apex between the
veins R5 and M1, a double lack spot between M1 and M3, the next [from the apex]
intervein plate contains a dot, the next one contains a round black spot. No
adroconial brand is seen. On the UPH there are four round black spots on an
ochre fulvous band going from the fore margin to the vein Cu2. The third spot
(counting from the fore margin) is noticeaby shifted outside. The underside is
analogously colored but the round color is darker to some extent, the band on
the FW is ochre-yellow, while the black dot at the apex is absent; on the HW the
spots are smaller and reside not on the band but in narrow ochre rings, inside
of which there is a white band split into fragments. The fringe is chequered. In
the genital apparatus the valva is narrow (the ratio of its maximal width to the
length being 1:5), slightly and gradually tapering to the apex; a half of its
dorsal edge bears teeth of various size (see Appendix). In a similar sp. E. ligea the valva is sharply (two- or three-fold) narrowed at
the apex forming a finger-like projection. A structure of the male genitalia
similar with that of E.
kosterini is found in an American sp. E.
vilderi Elwes, 1898 [an error in the original description, must be vidleri
Yu. K], well differing by the absence of postdiscal black ocelli on the HWs.
ALLOTYPE:
a female. FWl.: 20 mm. The wings are noticeably narrower than in the holotype.
The coloration and pattern are similar to those of the holotype, differing in
the following: on the UPF the black dot at the apex is absent, on the UPH the
black spots are smaller and there is only three of them (that colsest to the
fore margin is wanting); the double spot on the UPF, the same double spot and
the spot between Cu1 and Cu2 [on the UNF these words were accidentally
dropped in the printed version Yu. K.] , and three of the four spots on the
UNH have white nuclei. On the UNH the white stripe inside of the row of ocelli
is brighter and more clear-cut than in the holotype.
MATERIALS:
Holotype: ♂
20.07.1989, the Magadan Region, the Koni Peninsula, the lower reaches of
the Khindzha River, a herbaceous meadow on the lower terrace (O.E.Kosterin).
Allotype: ♀ 20-24.07.1989, the same locality. Paratypes: 2 ♂♂
2 ♀♀
20.07.1989, the same locality and date. The sp. is named by the surname
of Oleg Engelsovich Kosterin, a participant on numerous expeditions in
Siberia and the Far East, who collected the type series of butterflies of the
sp.
Origin:
Dnevnye babochki, Ekaterinburg, 1995,
p. 123.
NOTE:
A more detailed description of the habitat of these butterflies (referred to as Erebia
ligea [Linnaeus, 1758] group) on
the Koni Peninsula can be found in [Kosterin, 1993]. After the first description
the butterflies of this sp. were collected also at the Magadan City and the
settlement Karamken in the Magadan Region [Korshunov, 1996].
NOTE.
The entire puzzle began with an incorrect definition by P. Gorbunov of the
butterflies in the collection of V. V. Baglikov from the Southern Sikhote-Alin;
the author was on a hurry to publish a text regarding the supposed capture of kosterini
in a new site. I learned about this text incidentally only much later.
Nonetheless, I immediately doubted the existence of the special population of
this form in the Southern Sikhote-Alin, because I was previously familiar with
the details of the description of the ssp.
arsenjevi and realized that its
description unfortunately was not taken into account during the definition of kosterini.
It was also clear that arsenjevi
was not a ssp. of ligea,
but rather an entirely different butterfly. This conclusion was then reported to
V. Dubatolov. The further details on kosterini
and ligea became clear through
our collective efforts. A. Lvovsky helped us especially having sent the
necessary data about the specimens located in ZIN. A separate publication (in
Trans.lepid. Soc. Japan, v.49, 1998) was devoted to a description of the
lectotypes both of ajanensis and eumonia.
The separate sp. janensis was
first cited in [Elwes, 1881], then in [Graeser, 1888], a fact which every first-discoverer
should bear in mind.
Erebia aethiops
(Esper, 1777)
Many
aberrations of this sp. are known. In particular, the large form called uralensis Goltz,
1930, referred later to the Alpine ssp. sapaudia Fruhstorfer,
1917, has been designated later marked as an Uralian one, which
seems entirely not justified. It is impossible to call the Uralian butterflies
by this name, since there is already such a taxon of .
medusa.
In this connection we have introduced for the Uralian specimens another name, goltzi Korshunov,
1998. It differs from the eastern butterflies by a darker background
and by a redish band on the FW with a less noticeable tie. The UNH is almost
monochrome in males; the dusting in females is rather yellowish. Similar
butterflies were earlier unsuccessfully called altaica Goltz,
1930 (the name is invalid because of the homonymy). It was replaced
by us through transsibirica Korshunov,
1998. O. Staudinger described a male
with a wing span of 40 mm and a cosiderable reduction of the pattern similar to aethiopella
Staudinger, 1897 (ntei
is located in the Malkhanski Mt. Range in the Chikoi R. basin Yu.K.), which was corrected later as ab. ignota Higgins,
1930 because of the homonymy. Basically, small butterflies in the
populations in the S. Siberia are not rare, but it was impossible for us to
collect specimens identical to ignota.
Erebia rossi
Curtis in Ross, 1834
The
ssp. ro
Bremer, 1861
occurs in the much of the S. Siberia (TS
Jablonovy Mt. Range). The butterflies have an erubescent-chestnut-brown
ground color of the UNF (the Altaian specimens have no original features,
despite the claim of [P. Gorbunov, p. 125]!). The forms erda Sheljuzhko, 1924
in the E. Sayan, dzhelinde
Sheljuzhko, 1924 in the Dzhalindinski Goltsy in the Jankanski Mt. Range
(mountains of the Upper Priamurye) have also been described. Paul Ehrlich
[Ehrlich, 1952: 75-88] considers all these titles as synonyms of ero
in his comprehensive paper about the occurrence and the ssp. of rossi.
The butterflies found in the tundra and forest tundra of North Asia and in the
mountain communities of Yakutia and Chukotka are distinct. We have described
them as the ssp. subarctica.
Erebia rossi
subarctica Korshunov, 1996 (fig. 1,
2,
3,
4,
5,
6,
7,
8).
Translation
of the original description:
...The
butterflies from the zonal tundras and forest-tundras of N. Asia and from
mountains of Yakutia and Chukotka are different. They differ first of all by the
absence of a reddish-chestnut-brown color on the UNF in a majority of specimens.
Only some females have a brand of this color and few males have a group of such
scales at the ocelli. We describe them as Erebia
rossi subarctica Korshunov, ssp. n.
HOLOTYPE:
a male. FWl. 24 mm. The UPS is brown, there is a double fulvous spot at the FW
apex, containing two black dots, and a a hardly noticeable fulvous spot without
a dot at the Cu veins. The UPH has no dots. On the UNF there is a double fulvous
spot with two black dots centered with white ocelli, a tiny fulvous speck and a
small fulvous spot, with a white-centered black dot, between the branches of the
vein Cu. On the UNH there is an obscure band, with a white dot on its margin at
the cell apex, and a somewhat ligher band in the outer field. There is a flash
of reddish scales at the spots, noticeable only if to place the butterfly at
different angles to the light.
ALLOTYPE:
a female. FWl. 25 mm. The UPS is as in males, but the small spot at the Cu is
missing. The UPF has only two black dots with white pupils in a common fulvous
round spot at the apex. The UNH has the same bands as the male, the outer margin
of the outer one bearing two very small white spots.
Among
the paratypes about a half of specimens have 2-3 spots in addition to the double
apical one on the FW and 2-4 spots on the HW. On the UNH there may be fore white
dots. The imagines from the Maymecha River basin and from the more northern
Novaya River, on the Putorana Plateau, are peculiar: they are smaller, FWl.
being 19-21 mm, and have only the apical spots (the only female of 8 specimens
has spots also on the UPH), this population deserves further studies.
MATERIALS:
the holotype: ♂ 15.07.1968, the left bank of the Tanlovaya River, a tributary
of the Schuchya River, S Yamal (Korshunov). The allotype: ♀
14.07.1968, a tundra between the Tarseda River and the Schuchya River left bank,
S Yamal, Korshunov). Paratypes: 10 ♂♂ 7 ♀♀ 14-
30.07.1968, the Schuchya River basin (Yu. Korshunov); 2 ♂♂
28-29.06.1969, the Omolon River valley (N. Gomoyunova); 2 ♂♂
8.07.1976, the Yenisei River lower reaches, Chopko (Chernenko); 2
♂♂- 07.1981, Taymyr, Ary-Mas, Novaya (R. Polovinkina); 1 ♂ 5
♀♀ 28.07-8.08.1982, Putoran, Maymecha (I.S. Zakharzhevsky); 1
♂ 22.06.1984, the SutarKhayata mountain range (L. Popova); 14
♂♂ 1 ♀ 19.06- 14.07.1985, the Suntar-Khayata mountain
range, the Khandyga River headwaters (V. Dubatolov); 1 ♂ 18.06.1989,
Yakutia, the Kele River headwaters, a Carex/Eriophorum
tussocks (L. Popova); 2 ♂♂ 12-17.07.1993, the Chita Region,, 35
km SSE of Chara, Naminga (A. Streltsov).
Origin:
Dopolnenie 1, Novosibirsk, 1996, p.
33-34.
NOTE.
In the book of the Moscow collectors Butterflies of Russia, v. 1, 1997 the
ssp. subarctica
described by us is considered as a synonym of the American ssp. kuskoquima
Holland, 1931, which
occurs in the Kuskokvim R. basin. The UNF of this ssp. are almost
rubescent-chestnut, and the other features different from those butterflies in
the table 48, no. 1-3, all of which serves only to confuse the reader.
Our
name subarctica
is invalid because of the presence of Erebia
disa subarctica McDunnough, so we give this ssp. a new name Erebia rossi yamala Korshunov,
nom.n.
Erebia semo
Grum-Grshimailo, 1899 (= fasciata auct., nec Butler, 1866)
The
sp. semo
is widespread in Siberia to the north as far as the Chukotski peninsula
inclusive. The ssp. avinoffi Holland, 1930
occurs on Wrangel Island; the butterflies have reduced chestnut spots on the UPS
of w. The ssp. sachaensis Dubatolov,
1992, stat.n. is found in Suntar-Khayata, Cherski and the Esop
Mountain Ranges. The females have brown bands (the first-description writes that
the bands are as in semo!
― Yu. K.); the males have barely peceptible bands or one-colored
wings. The scales on the whole and the androconial ones in particular are
similar to semo, but they are not identical. The same could be said about
male genitalia. It is quite possible, that sachaensis
is a bona sp.!
rebia semo sachaensis Dubatolov, 1992 (fig. 1,
2, 3,
4).
Translation
of the original description:
rebia [magdalena]
sachaensis Dubatolov, ssp. n.
Materials
[in ISEA, Novosibirsk]: holotype ♂, Yakutia, 180 km ENE of settlement
Khandyga, the upper flow of the East Khandyga River, 248-249 km of the road
Khandyga-Magadan, the Setarym River right bank, a south-western slope, 1500 m
above sea level, at a scree, 28th June 1985 (Dubatolov). Paratypes: 3
♀♀, the same locality, 1500-1600 m, mountain rocky tundra,
11.07.1985 (Dubatolov); 1♂, the same locality, 28.06.1985
(Barkalov); ♂, the Vostochnaya Khandyga River headwaters, 232 km of
the road Khandyga-Magadan, the Baranya Mt., 1500 m above sea level, a crest, 7th
July 1985 (Barkalov); a female, 10 km of settlement Kyumyuba, 336.5 km of the
road Khandyga-Magadan, 18th July 1985 (Dubatolov); a male, the
Verkhoyanskii Mt. Range, the Kele River headwaters, the Gnas River, a left
tributary of the Kyunkyunyur River, 8th July 1989 (Vinokurov); a
male, the same label without date; a male, the Cherskii Mt. Range, the Burkat
Pass between the headwaters of the rivers Myuryule and Inyali, 10yj July 1990
(Savin); a female, the same locality, the Burkat spring, a Myuryule River
tributary, 11th July 1990 (Zinchenko).
Male:
FWl.: 25-27.5 mm. FW widely rounded at apex, black-brown with a brownish-red
diffuse spot on outer area between the veins M3―Cu1, Cu1―Cu2 and
below Cu2. This spot may be large, up to 5 mm wide, or much reduced but always
present. The UPH are evenly dark-brown. The UNF have a rectangular
dark-brownish-red spot on the outer part between the middle of space
M3―Cu1 and the fold A. Sometimes basally of this area there is a
triangular browinish-red area with an apex directed to wing base, sometimes it
is reduced but in this case some brownish-red scales remain in this place.
The
HW either evenly black or with a lighter base and a light band, as in E.
fasciata semo Gr.-Gr. Androconial scales present on the FW, their shape
is as in E. magdalena Strecker.
Female:
FWl.: 24-25 mm. FW above blackish-brown with a reddish-fulvous area bordered
with middle of cell, vein M2 and fold A. This area is clearly cut through with
black veins. The UPH are evenly blackish-brown. UNF are as UPF, the HW are like
in E.
fasciata semo.
Systematic notes. In the wing coloration the new taxon resembles E.
fasciata semo from which it differs with rounded FW, a diffuse fulvous
band and also a presence of androconial scales. The Siberian ssp. differs from
the Alaskan E. magdalena mckinlyensis Gunder (We have seen the only one
picture of a female of this taxon in [Emmel, 1975: 10, fig. 4]) by a smaller
size of the fulvous spot on the UPF and darkened veins.
Biology: The butterflies fly on stone screes, rocky slopes, and also on stone
mountain tundra on the altitudes of 1500-1600
Origin:
Vestnik zoologii, Kiev, 1992, No. 6,
pp. 41-42.
NOTE.
The authors of the Moscow collectors Butterflies
of Russia, v. 1, 1997 treat semo
as if it were an individual variation, but the basis of this assertion is
simply unintelligible. The baseless desire to demole this or that taxon is
has no basis in fact and serves no purpose.
Erebia ola
Korshunov, 1995 (fig. 1,
2, 3,
4, 5,
6, 7,
8).
Translation
of the original description:
Specimens
from the Magadan Region are characterized by a complete vanishing of
chestnut-colored spots on the UPS in males and their reduction in females. Below
we describe them as a new ssp. The shape of the valva in the male genitalia in sachaensis
and ola
is very variable, that is illustrated by the drawings of the genitalia of the
holotypes (see Appendix). In general, the genitalia of these taxa have no
significant differences between each other, as well as from E.
semo and E. erinnyn. Erebia
[mckinleyensis] ola
Korshunov, ssp. n. [this
combination is an invention by Mr. P. Gorbunov!!- Yu.K.].
HOLOTYPE:
a male. FWl. 25 mm. The UPS is evenly soot-colored. The underside is of the same
color but on the FW, at the cell, there are groups of reddish scales forming a
hardly detectable sinking spot. The light scales are scattered over the UNH
where no bands are seen. In the genitalia the valva has a rounded apex.
ALLOTYPE:
a female. FWl. 26 mm. The UPS is black with a brown tint. On the FW, at the
cell, there are reddis elongate streaks between the veins. On the UNF the
corresponding reddish spots are bordered with dark scales. The UNH is not only
mottled, as in males, but have also distinct traces of the postdiscal band, the
margin of which is most clearly seen in the middle part of the outer field.
MATERIALS:
the holotype: ♂ 7.07.1991, the Magadan Region, the headwaters of the Ola
River, the surroundings of the mountain Nukh (1828 m) (V.V.Palekha leg.); the
allotype: ♀ 7.07.1991, the same locality; paratypes: ♂
07.1991, the same locality; 2 males 1 ♀ 13.07.1993, the same locality;
♂
6.07.1964, the Omsukchanski mountain range, at the setlement Golimyy, 6th
July 1964 (A.V.Tsvetaev).
The
attribution of this taxon to E. mckinleyensis as a subcpecies, made by P. Gorbunov in a
published version of the book [Korshunov, Gorbunov, 1995], is erroneous. I
consider it to be a bona sp. due to
the complex of characters concerning the general appearance, pattern, the
details of the genitalia structure [Korshunov, 1996]. The new sp. has a certain
similarity to E. erinnyn Staudinger, 1894, but this sp. of the E Sayan lacks
androconia resembling E.
ola, their genitalia structure have substantial differences, in
particular, the end of the valva is twice as thick in E. ola, in general the valva is thicker and resembles that in E.
mckinleyensis.
Well-developed spots in males and, to some extent, in females is another trait
differing E. erinnyn from E.
ola. I am not satisfied with the original description of the holotype
and would suggest a better version [Korshunov, 1996].
Origin:
Dnevnye babochki, Ekaterinburg, 1995,
p. 127; Dopolnenie 1, Novosibirsk, 1996, p. 35.
NOTES
REGARDING CLASSIFICATION. The new sp. bears a definite similarity to E. erinnyn Staudinger,
1894, but the sp. from the E. Sajan has no androconia similar to olas
ones; there are noticeable genital differences, specifically, the extremity of
the valvae is nearly twice as wider, the same valvae are wider as a whole and
similar to the valvae of E.
mckinleyensis (see the drawings).
The explicit development of the spots in the males and, to a greater degree, in
the females of erinnyn
is an additional difference from ola.
Erebia erinnyn Staudinger,
1894 (= sajanensis Korshunov,
1972)
In
an American publication [Scott, 1986] erinnyn
is equated with the American magdalena
and mckinleyensis.
We have previously shown that they differ markedly in various details. Here it
is necessary to note that these American sp. are trophically connected with Luzula
spicata and Carex
atrata. The eggs are laid one by one on the stones near the bushes of
Luzula. A dark green
caterpillar with thick short hair gnaws round the ends of the leaves. The
trophic attachments of erinnyn await classification, but it is necessary to remember, that L.
spicata and C.
atrata grow in the E. Sayan in moss-lichen and petrous tundras. The
reader should bear all the above in mind.
Erebia jakuta
Dubatolov, 1992 (fig. 1, 2,
3, 4).
In
our book of 1995 jakuta
belongs to E. occulta R. et K., 1983. Further comparison of the
butterflies, however, in size and pattern, as well as details of genitalia
showed that occulta is a separate sp. that occurs only in N. America.
Translation
of the original description:
Erebia
anyuica jakuta Dubatolov, ssp. n.
Materials.
Holotype, a male, Yakutia, 180 km ENE of settlement Khandyga, 232 km of the road
Khandyga-Magadan, the mountain crest, 15th June 1985 (Dubatolov).
Paratypes: 7 males, 4 females, the same label, 7.07.1984 (Popova), 15.06-
11.07.1985 (Dubatolov, Barkalov); a female, 248 km of the road Khandyga-Magadan,
the Seratym River right bank, a plateau, 1550 m, 28.06.1985 (Dubatolov); 8
males, E Yakutia, 100 km NE of Ust-Neram the Siyap River, 20-30 km upstream
of the mouth, 1-5.07.1990 (Zinchenko, Savin); 4 males, 1 female, 170 km NE
Ust-Nera, the Myuryule, the brook and pass Burkat, 10-11.07.1990 (Zinchenko,
Savin)
Male.
FWl. 20-24 mm. UPS are dark-brown, the FW along the fore margin with 4 fulvous
round spots centered with black dots of variable size. These dots of two fore
spots are always stretched transversally cross the wing. The HW wings between
veins Cu2 and M3 and in front of M3 with the same spots are centered with round
black spots. The UNF pattern is the same as on the UPF but tyhe fulvous spots
are fused into a band, between the veins M1 and A. This band widens basally to
the large triangular spot, the apex of which is directed towards the wing base.
The black dots on this band can disappear. The UNH has a pattern as in E.
dabanensis but more reduced. Genitalia are as in a. occulta Roos et Kimmich,
1983, the valvae are without a heel-like projection.
Female.
FWl is 21-23 mm. The wing pattern as in male, but black dots are larger,
sometimes these FW dots are more than 1 mm in diameter.
Systematic notes. Basing on the male genitalia structure (the valvae
without a heel-like projection, well seen in E. dabanensis, and the
disposition of the teeth on the valva apex ― they extend noticeably from
the dorsal margin towards the outer margin), anyuica
Kurentz. and occulfa
Roos et Kimmich (= E.
phellea Philip et Troubridge) are conspecific. The new ssp., different
from a.
anyuica, has a well developed pattern, conspicuous even in the
melanistic specimens. The butterflies of the American ssp. a. occulta are much smaller than the nominotypical (FWl is
16.1-20.2 mm), the wing pattern is less expresed (the diameter of the spots on
the FW is 1.0-1.5 mm, instead of the 1.5-3.5 mm in a new ssp.).
Origin:
Vestnik zoologii, Kiev, 1992, No. 6,
p. 44.
In
Asia we found out such ssp. of jakuta. E.Sayan
the ssp. iltshira
Belik, 1996, stat. n. (T.S.
Kitojskie Goltzy). These butterflies have black dots on the UNF.
Erebia jakuta
iltschira Belik, 1996 (fig. 1,
2)
Original
description:
Erebia anyuica iltshira,
ssp. n.
Erebia
kozhantschikovii {sic} ab. rubescens
Warren, 1930, infrasubspecific name; Erebia
dabanensis {sic} ab. rubescens
Warren, 1936, infrasubspecific name.
Type
material. Holotype: male: Russia, Vostochnyy Sayan mountains, khrebet (mountain
range) Kitoiskiye Goltsy, river Kitoi upper stream, vicinity of the lake
llchir, 2250 m, 26.VI. 1994. A. G. Belik leg. Paratypes: 42 males, 9 females.
same locality as holotype, 26.VI and 30.VI. 1994, A. G. Belik and E, G. Belik
leg.
Description.
Male
(plate I, figs 1-4), average FWl. 22.1 mm (range 21.0 24.0 mm). FWl of the
holotype 22.5 mm.
UPF:
ground color dark brown with a golden fringe. Four submarginal spots between M1
and Cu2;. colored from reddish-brown to ochre-orange, with diffuse outline
especially towards the wing base, often forming a band. When separated, these
spots look like diffuse oval ocelli with minute black pupils. There is sometimes
an additional submarginal spot between Rs―M. The fringe monochrome as in
the FW.
UPH:
ground color as in the FW. Between M2―Cu2 are three reddish-brown to
ochre-orange submarginal spots with slightly diffuse outlines. Now and then
there are additional spots between M1―M2 and Cu2―2A. The submarginal
spots are sometimes centred with minute black dots. The fringe is concolorous
with the wing.
UNF:
the ground color is dark-brown. Wide submarginal band is usually reddish-brown
to ochre-brown, generally darker than the corresponding spots on the upperside.
The space between the submarginal band and the wing base are usually with
reddish tinge because of diffuse reddish scales, which are more numerous near
the inner margin of the submarginal band and cause a more prominent reddish
color to this area. Thus the border between the submarginal band and the darker
postdiscal area is often not sharp. Black dots inside the submarginal band
corresponding to those on the upperside that centre the submarginal spots, are
somewhat larger than on the upperside.
UNH:
the ground color is blackish-brown, very dark. Sometimes a submarginal band of a
slightly paler color is visible, especially in tn oblique light. Usually three
reddish-brown submarginal ocelli between M^-Cu^ contain black pupils of larger size than in
upperside submarginal spots. The fringe is concolorous with the wing.
Genitalia
(Fig. 1): of similar structure to other sp. of the Erebia
dabanensis complex. The shape of valva in Erebia anyuica varies within broad limits (figs 1, 2. 5). being
somewhat similar to that of E.
kozhantschikovi Sheljuzhko, 1925
(as in fig. 5. though Kurentzovs figure seems somewhat schematic) or of E. dahanensis Erschoff,
1871 (as in fig. 2). The main characteristic feature of E.
anyuica are male genitalia, which differ it from the related sp.: the
presence of an additional row of spines on the outer surface of the dorsal side
of the valva in its distal half. Thus the valva of E. anyuica has 3 rows of spines, while the valva of both E.
dahanensis (fig. 3) and E.
kozhantschikovi has 1-2 rows of spines only along the costal edge in the
distal half and has no spines on the outer surface of the valva.
It
is surprising that Warren [Warren1930; 1936] has overlooked this fact and never
recognized his specimen as belonging to a new sp. He considered it only as an
aberration. First of E. kozhantschikovi and then of E. dabanensis: though on the
valva of his specimen the additional rows of spines are much reduced, they are
visible as two short additional rows in the extreme distal part of its outer
surface [Warren, 1936: pi. 42, fig. 384].
Female
[plate I, figs 5-8: the average fore-wing length 22.5 mm (range 22.0-23.0 mm).
UPF: ground color paler than in male, submarginal ochre-brown spots often united
into a band 3-5 mm wide. 3-4 black dots inside this band are larger than in
male, up to 0.5 mm in diameter.
UPH:
as in male, but paler in color. 3 to 5 submarginal reddish-brown or ochre-brown
ocelli larger than in male.
UNF:
as in male but paler. Submarginal band usually ochre-brown. Area between the
inner margin of the band and the base of wing are usually hushed with color,
slightly darker than that of band. Submarginal black dots are often centred with
white nuclei.
UNH:
general pattern as in male, but much more visible. The wing is usually heavily
dusted with pearly gray scales that produce a silvery-grayish tint. This dusting
shade wing pattern that is often well visible as a dark heavy jagged marginal
band and a dark basal-discal area with jagged outer margin. Submarginal ochre
ocelli are prominent but not very conspicuous on the silvery-grayish band.
Origin:
Phegea 24(4) (1.XII.1996): 157-159.
In
Pribaikalye and Zabaikalye the ssp. sokhondinka Dubatolov et Zintchenko, 1995, stat.n.
Erebia jakuta sokhondinka
Dubatolov et Zintchenko, 1995 (fig. 1,
2, 3)
Translation
of the original description:
Butterflies
from the Chita Distr., described below, are much larger than E. o. occulta (FWl. being
19-21 mm) and significantly larger than E.
o. jakuta (FWl. 20-24 mm). They well differ from both known ssp. by more
developed fulvous marking on the wings, often forming a contiguous band on the
FW. There is also a significant difference in the length of the dentate part of
the valva. Its ratio to the total length of the valva in E.
o. jacuta is 52-67% (with
the mean 59,5%), the samlpe size being 24, in new ssp. it is 56-67,4 (with the
mean 64%), the sample size being 20. The butterflies from the East Sayan and
Pribaikalye are close to the ssp. being described.
Erebia
[occulta] sokhondinca
Dubatolov et Zinchenko, ssp. n.
Male:
FWl. is 23 mm in the holotype, 23-26 mm in paratypes. The UPS is dark-brown, the
FW bears 4 large pale-fulvous spots along the outer margin, usually fused into a
band. They are centered with black dots elongate lengthwise with respect to the
wing. The HW has 3-4 round spots at the outer margin, often centered with black
dots. The pattern of the UNF is the same as on the upperside, but the fulvous
spots are always fused into a bright band. This band can extend towards the wing
base. The UNH as in E. o. jakuta.
Female:
FWl. 23.5-26 mm. The pattern is as in males, but the black dots are larger, up
to 2 mm in diameter.
MATERIALS:
The holotype ♂, 23.06 1991, the Chita Region, the Sokhondinskii nature
reserve, the Verkhnii Bukukun River (Dubatolov). Paratypes: 35 ♂♂ 9
♀♀, the same locality and the mountain Tsagan-Ula, 19.06 6.07
1991 (Dubatolov, Zinchenko and others).
Origin:
Dnevnye babochki, Ekaterinburg, 1995,
p. 128.
The
first fixation of this butterfly in the Baikalski Mt. Range belongs to O. K.
Gusev. This unique specimen is represented in fig. 2 without any exact
definition [Korshunov, 1970: 189-190]. The specimens from Udokan were described
as the ssp. udokanika
Streltzov, 1998 [Streltzov, 1998:
1-4, figs. 1-3], since they are practically identical to jakuta. The nominotypical ssp.
jakuta differs from iltshira
and sokhondinca
by the more or less roundish erubescent spots and prominent black points. The
genitalia of the ssp., as is evident in the tables, are similar, but their
difference from occulta is obvious and requires no explanation.
Erebia theano
(Tauscher, 1806)
We
have described both ssp. tshugunovi and ssp.
shoria.
The ssp. tshugunovi from Salair differs by brick-red coloration of spots
in the UPS, all spots are realatively small, especially on the UNH.
Erebia theano
tshugunovi Korshunov et Ivonin, 1995 (fig. 1,
2,
3,
4,
5,
6).
The
translation of the original description:
...Below
we describe two ssp. from the Salairski Upland and the Kuznetski Alatau Mts. The
former is characterized by a brownish color of spots on the UPS. The butterflies
from the Kuznetski Alatau, West Sayan and Tuva are characterized by a general
reduction of ochre-brown spots not forming contiguous bands.
HOLOTYPE:
a male. FWl.: 18 mm. The UPS is dark-brown; the postdiscal spots and the spot in
the cell are brick-red.The underside is the same as the upperside, but with a
grayish tint, which is more distinct along the wing margin; the fulvous
postdiscal spots on the HW consist of four hardly noticeable dashes. The fringe
is white with dark markings.
ALLOTYPE:
a female. FWl.: 20 mm. The UPS as in the male. The gray tint on the underside,
especially on the HW, is much more pronounced. The yellowish postdiscal spots on
the UNH are smaller than on the FW (where they are fulvous) and smaller than in
the male; the basal spots on the HW are small and hardly noticeable, as in the
male.
MATERIALS:
the holotype: ♂, 21.06.1994, the Novosibirsk Region, the Kiternya river
valley, a damp meadow (V.V. Ivonin); the allotype: ♀ ― 20.06.1994,
the Novosibirsk region, the Bolshoy Elbash river valley, a damp meadow (V.V.
Ivonin). The paratypes: 11 ♂♂ 8 ♀♀ 20.06.1994, the
same locality; 5 ♂♂ 21.06.1994, the same locality,
♀♀ 24.06.1994, the same locality; 5 ♂♂ ―
21.06.1994, the Novosibirsk Region, the village Novoselovo [this is a mistake
should be Novososedovo, ― Yu.K.],
a forest opening (V. V. Ivonin); 1 ♂ 24.06.1949 the Novokuznetsk
city surroundings, the village Torgay (A.E. Shtandel); 2 ♂♂ the
Tomsk region, the village Nizhnie Sokoly, a bogged meadow (G.S. Zolotarenko)
The
ssp. is named in honour of Sergey Mikhaylovich Chugunov, a doctor of medicine at
the Tomsk University, who in 1891-1920 comipled several annotated lists of
Lepidoptera for various regions of Siberia.
The
Kuznetskian ssp. differs by a hardly perceptable spot in the discoidal cell on
the UPF and bright ochre-yellow coloration of the band spots.
Erebia theano
shoria Korshunov et Ivonin, 1995 (fig. 1,
2,
3)
Translation
of the original description:
Erebia theano
shoria Korshunov et Ivonin, ssp. n.
HOLOTYPE:
a male. FWl.: 19 mm. The UPS is brown with the postdiscal bands composed of
ochre-yellow spots and a hardly noticeable spot in the cell. On the UNS all the
spots are bright and large, including that in the cell of the FW and the basal
spots on the HW; the gray suffusion is well expressed in the basal area. The
fringe is grayish, chequered.
ALLOTYPE:
a female. FWl.: 19 mm. The UPS as in the male, the spot in the cell is well
expressed; on the UNS all the spots are bright and large; on the HW the basal
spots and gray suffusion are well expressed. The specimens of the type series
vary in size of the lighter spots, especially of the basal spots on the UNH. The
fringe is brownish in some males.
MATERIALS:
the holotype: ♂ 5.07.1969, the Tuzukhsu river valley in the Tom
river headwaters 1969 (Yu.P. Korshunov). Allotype: ♀ ― 13.07.1992,
the Gornaya Shoria mountain area, the Sheregesh settlement environs, the
mountain Pustag, 1300 m [above sea level] (V.V. Ivonin). The paratypes: 21
♂♂ 5.07.1969, Gornaya Shoria, the Tuzukhsu valley; 13
♂♂ 9 ♀♀ 28.07.1969, the same locality; 86
♂♂ 28 ♀♀ 24.06-27.07.1969, the Askiz River basin,
Birikchul (Yu.P. Korshunov); 23 ♂♂ 17 and 24.07.1969,
Khakasia, the Baza river headwaters, (Yu. P. Korshunov); 4 ♂♂ 2
♀♀ 13.07.1992, the Sheregesh settlement, the mountain Pustag
(V.V.Ivonin)
Origin:
Dnevnye babochki, Ekaterinburg, 1995,
p. 130.
The
Altaian butterflies have large spots in the central cell, the band spots color
is reddish-yellow or yellow.
The
exact sample site of the ssp. theano in the Altai is not known. One can only hypothesize that
it can be located in the western part of the Altai Mts. From the C. Altai
altitudes higher than 1800 m the ssp.
lederi
A. Bang-Haas, 1906 (Iris, XIX: 127-128) was described; the butterflies
have wide ochre spots on the exterior margin of the wings. The background of the
UNS is lighter in color and more yellow than the color of theano.
A. Bang-Haas indicates the wing-span 38- 40 mm (males) in theano and 30-32 mm
(males), 35-36 mm (females) in lederi.
In the book by Warren lederi
Goltz, 1930 is cited, but it is a
synonymous taxon of lederi A. Bang-Haas
(Goltz cites this taxon as lederi
A. Bang-Haas for no apparent reason
in the Supplement to Seitz, i.l.! ― Yu.
K..). Furthermore, Warren [Warren, 1936] considers it as a ssp.
Nevertheless, the drawings of lederi
in the Supplement to Seitz (v.10, A, No.3) and in the book by Warren do not
coincide. Perhaps, this is simple due to the variability, and there is no basis
of a ssp.; it is necessary to verify the primary evidence. The ssp. approximata, described by Warren on the sole specimen from the higher
elevations of the Korgonski Mt. Range, Altai, is assigned in the book by
Lukhtanovs [Lukhtanov & Lukhtanov ,1994] for an unknown reason to the Altai
foothills and their vicinity. Our available specimens from the Mountainous
Kolyvan etc., for example, are similar to the typical theano. In Warrens
photos of the specimen 7 of approximata,
the exterior features and the details of genitalia do not coincide with those of
tshugunovi.
Erebia dabanensis
Ershov, 1871 (= tundra Staudinger, 1887).
The
nominotypical ssp. occurs in Pribaikalye and the Stanovoj Upland; precisely
these butterflies can have oblong ocelli. The specimen caught in the Providenie
Bay on Juny, 27 1900 by A. N. Akifieva stands apart; it is marked as tschuktscha Herz,
1903. It is small, partly resembles the American youngi
with an untrained eye, and is not similar to troubridgei. Its pattern
is similar to this of dabanensis.
It is also similar to the ssp. olshvangi
that is smaller, darker and has an noticeable reduction of the brown postdiscal
spots in comparison with the type. But the specimen from Providenie Bay
doesnt coincide with the Uralian one in genitalia: the valvae have an
explicit jagged heel similar to the nominotypical dabanensis:
the heel of the Uralian butterflies is rather oblong (in the book of 1995
there is an error on the page 188, fig. 270). In any event it turns out that on
the edge of the northern Asia occur small butterflies of the type dabanensis that may be
treated as a special sp., which should be defined as tschuktscha Herz, 1903; if
new evidence arises, the limits of the genital variability can be specified
The
ssp. dabanensis is characterised by a continuous brown postdiscal band in
females and oblong black blind ocelli in both sexes.
Below
follow descriptions of two ssp.
Erebia
dabanensis troubridgei Dubatolov, 1992 (fig. 1, 2,
3).
Translation
of original description:
Materials
[in ISEA, Novosibirsk]: the holotype ♂, Yakutia, 180 km NEE of
settlement Khandyga, the Vostochnaya Khandyga River headwaters, 232nd
kilometre of the road Khandyga-Magadan, 1st July 1985 (Dubatolov)].
Paratypes: 55 ♂♂ 6 ♀♀, the same locality, 16th
June 14th July 1985 (Dubatolov); 25 ♂♂ 6
♀♀, the same locality, 248-249th km of the road
Khandyga-Magadan, the Setarym River, 28th June and 11th
July 1985 (Dubatolov); ♂, the same locality, 250th km of the
road Khandyga-Magadan, the Shagali River, 20th June 1984 (Popova);
♂, the Cherskogo Mt. Range, the Inyali River 50 km upstream of its mouth,
15th July 1987 (Dorofeev); ♂, the Cherskogo Mt. Range, the
Ytabyt-Yuryakh River, 10-40 km upstream of its mouth, 4-9th July 1987
(Dorofeev).
Male.
FWl. 20-25 mm. The UPS are dark-brown, at outer margin there are usually four
rounded reddish-fulvous spots centered with black dots slightly stretched out
transversally; sometimes an additional fulvous spot is developed, blind or with
a black dot above the vein M1. The spots in this row are always rounded, well
separated from each other. The patterns on
the UNS and also on the UPH are as in nominotypical ssp.
Female.
FWl. 20-23 mm. The UPS are light-grayish-brown with a pattern as in male, the
spots in the outer zone are well separated also.
Systematic notes. The new ssp. is in general larger than the
nominotypical E. d. dabanensis Ersch. (= E.
tundra Stgr.), described from the Khamar-Daban Mts. (the FWl. being in
males 19.5-21, in females FWL. 18-20 mm). The spots in the outer area on the UPF
are always rounded, well separated even in females. In E. d. dabanensis
the spots are larger, usually, in females always, form a contiguous fulvous band
with four black spots; this band is divided by narrow black veins only.
Biology. This butterflie is common in the mountains of Yakutia, it inhabits open
larch forests, valley meadows, bogs; in the mountain tundra and stony screes it
flies together with a close Holarctic sp. Erebia occulta.
Origin:
Vestnik zoologii, Kiev, 1992, No. 6,
pp.43-44.
Erebia dabanensis
olshvangi P. Gorbunov, 1995 (fig. 1, 2,
3).
Translation
of the orininal description:
.....The
below described butterflies from the Polar Urals differ from other ssp. by a
darker brown-black ground color and the most substantial reduction of brown
postdiscal spots.
Erebia
dabanensis olshvangi
P.Gorbunov, ssp. n.
HOLOTYPE:
a male. FWl.: 20 mm. Both sides of the FW are dark-brown with four separate,
diffuse on the underside, fulvous-brown spots centered with small black dots;
both sides of the HW have three such spots; on the UNH the postdiscal area is to
some extent lightened due to grayish scales; the genitalia are ventralagous to
those of other ssp.
ALLOTYPE:
a female. FWl.: 20 mm. The UPS is brown, the underside grayish-brown; on the
UNH a light-gray postdiscal band is clearly expressed; the fulvous-brown pattern
is similar with that of the holotype. Generally the new ssp. is very variable.
The length of the FW ranges between 17 and 22 mm. In males the fulvous spots on
the upperside of both wings in 5-10 % of individuals are absent. On the UNF
they, however, never disappear entirely, on the contrary, in some specimens
(15-20 % of cases) they are fused into a common band, but its width, as
different from E.
dabanensis troubridgei, as a rule does not exceed 2 mm.
MATERIALS:
the holotype: ♂ 20.07.1992, Polar Urals, the railway station Krasnyy
Kamen, the mountain Slantsevaya, 300- 400 m above sea level, a montane
lichen-fruticulose tundra (P. Yu. Gorbunov leg.) The allotype 10.07.1998,
the same locality (V. N. Olshvang leg.) Paratypes: 7 ♂♂4
♀♀ ─ 10- 12.07.1998, the same locality; a male 5.07.1990
─ the same locality; 4 ♂♂ 3 ♀♀ ─
18-28.07.1992, the same locality.
The
ssp. is named in honour of Vladimir Nikolaevich Olschwang, a researcher at IEPA
UrD RAS [Institute of Ecology of Plants and Animals of Ural Division of Russian
Academy of Sciences] , an entomologist who for a long time investigated Polar
Ural and collected a part of the type series.
Origin:
Dnevnye babochki, Ekaterinburg, 1995,
p.129.
NOTE.
In some American works the continental dabanensis
is cited as Erebia
dabanensis youngi
Holland, 1900, and some
other works cite it as a separate sp. The review by [Troubridge & Philip,
1982 [83]: 107-146] justifies the independence of youngi.
Erebia callias
Edwards, 1871
The
ssp. altajana Staudinger,
1901 flies in the Altai and W. Sayan. In the Sayan Mts., Tuva and
Pribaikalye we find the similar ssp. simulata
Warren, 1933, whose butterflies have a lighter the coloring of the UNS. A ssp. tsherskiensis was
described from the basin of Indigirka.
Erebia callias
tsherskiensis Dubatolov, 1992 (fig. 1,
2,
3,
4,
5,
6,
7,
8,
9).
Translation
of original description:
Materials:
the holotype [in ISEA, Novosibirsk], ♂, East Yakutia, 185 km NW of
Ust-Nera, the Burkat Spring source, the right source of the Myuryule River,
1100-1200 m above sea level, 20thJuly 1987 (Dorofeev, expedition of the L.
Starikovskiis group). Paratypes: 23 ♂♂, 4 ♀♀, the
Myuryule River upper reaches, 13-16th July 1990 (Zinchenko); 18
♂♂, the Inyali River valley, 60-140 km upstream of its mouth, 18, 20th
June 1973 (Vinokurov). 9-10th July 1990 (Zinchenko, Savin, Popov).
Male.
FWl. 15-19 mm. The FW is doted at the apex. Two apical ocelli vary in size,
usually small, black, centered with white dots, always well separated from each
other and distinctly shifted to wing apex; they are situated on a
dark-reddish-brown area bordered by a discoidal vein and the vein Cu1. On the HW
there may be developed up to 4 reddish spots in the outer area, usually centered
with black dots. The UNS are as in E. c. simulata Wrn., but the HW are evenly colored, with specks,
while the apical ocelli on the FW are well separated. The valvae are short,
wide, without a heel-like ledge, teeth on their apical parts are dense. Female.
FWl. 15.5-18 mm. Wing pattern like in male, FW more rounded, ground color
slightly lighter. The fringes are different from males ones, they are lighter
than the ground color, some chequered. The pattern of the UNH is more mottled,
sometimes central band is weakly expressed and two thornal ocelli are hardly
noticeable.
Systematic notes: Hitherto 4 ssp. of E. callias were known: E.
c. callias Edw. ― the mountains of Colorado, North America; E.
c. sibirica Stgr. ― the Tarbagatai Mts.; E. c. altajana Stgr.
― the Altai; E. c. simulata Wrn. the Sayans, North Mongolia, the
Khamar-Daban Mts. [Warren, 1936; Lukhtanov, 1987]. Our specimens were collected
several thousand kilometres apart of all the known ssp.; they differ with the
pointed FW, well separated ocelli distinctly shifted to apex, and also by the
genitalia structure ― the valva is wide, the ration of its width in the
narrowest point to its length being 0.23, i.e. larger than in other ssp.; the
teeth on the apical part are densely set, as different from other Siberian ssp.
Origin:
Vestnik zoologii, Kiev, 1992, No. 6,
pp. 42-43.
Erebia fletcheri
Elwes, 1899
Erebia
fletcheri chorymensis
Korshunov, 1995 (fig. 1,
2, 3,
4,
5,
6)
Translation
of the original description:
...Below
Erebia
fletcheri chorymensis Korshunov, ssp. n. is described from the Kuznetski
Alatau Mts.
HOLOTYPE:
a female. FWl.: 22 mm. Both wing sides are brown; the FW has a wide
fulvous-brown band, slightly tapering to the ventral angle, with four oval black
spots about 1 mm long; On the HW three [the words of four was erroneously
omited here ― Yu.K.] fulvous
spots contain black dots, as well as an additional fifth small spot in front of
the ventral vein.
ALLOTYPE:
a male. FWl.: 22 mm. The postdiscal band on the UPF is darker and bears smaller
black dots. The paratypes have four of five fulvous spots on the HW, of which
one (in one specimen), three, four, or five being centred with black dots. In
the nominotypical ssp. (the holotype being a female) the black spots on the FW
are round and almost twice as large, the second on thr UNF being almost thrice
as large. Besides, in the butterflies from the Altai there is a tiny black spot
at the apex which is not situated in the same row with others, which is absent
in all the specimens from the Kuznetski Alatau. The HW in the nominotypical ssp.
has four roundish reddish-fulvous spots, the two of which at the ventral angle
being centered with black dots.
MATERIALS:
the holotype: ♀♀ 12.07.1975, the Kuznetskiy Alatau, the environs
of the mountain Bobrovaya (Yu.Korshunov); the allotype: ♂♂ the
same locality and date; paratypes: 5 ♀ 1 ♀ 9.07.1975, the same
locality; 4 ♂♂ 2 ♀♀ 12.07.1975, the same locality; 4
♂♂ 7.07.1975, the Kuznetskii Alatau, the Sarala river valley; 4
♂♂ 25- 26.06.1978, the same locality.The name is derived from
the Khakas word khorym a stone scree.
Origin:
Dnevnye baboschki, Ekaterinburg, 1995,
p. 132.
rebia fletcheri chajataensis Dubatolov, 1992 (fig. 1, 2,
3)
The
ssp. chajatensis
has been described for the Suntar-Khayata Mt. Range.
Translation
of the original description:
Materials:
the holotype [in ISEA, Novosibirsk], ♀ Yakutia, 180 km NEE of the
settlement Khandyga, the East Khandyga headwaters, 232nd kilometre of
the road Khandyga-Magadan, 23rd July 1985 (Dubatolov). paratypes: 8
♂♂, 3 ♀♀ the same locality, 14th June ―
12th July 1985 (Dubatolov); ♂, 248-249 km of the road Khandyga
―Magadan, the Seratym River, 28th June 1985 (Dubatolov).
Female.
FWl. 21.5-24 mm. The UPS dark-brown, fore ones with an ochre band in the outer
area. On this band, between the veins M1 and Cu2, there are 4 large oval black
spots, so that a distance between band border and spots is 2-3 times less than
spot diameter. Sometimes an additional small black dot is developed above the
vein M1.The UNS as in the nominotypical ssp.
Male.
FWl. 20-23 mm. Pattern on the FW is as in female but the band in the outer zone
of the FW is reddish-brown, further narrower than in females, while in one of
the paratypes it is split into separate spots, as in E. dabanensis troubridgei.
Systematic notes. The new ssp. is characterized by a more narrow band
on the FW and larger spots on it, so that the distance from this and the margin
to the spots is much less than the spot diameter in the nominotypical ssp. E.
f. fletcheri Elwes [Warren, 1936: pl. 101, fig. 1587, 1592] described on
a single female from the Altai, and also in other specimens from the western
mountains of S. Siberia, the fulvous band is wide, at least the distance between
its margin to the black spots is greater than the diameter of the spots
themselves.
BIOLOGY.
Imagines are rare, met with mostly on stony mountain slopes, less frequently in
the mountain tundra.
Origin:
Vestnik zoologii, Kuev, 1992, No. 6,
p. 43.
Erebia pandrose
(Borkhausen, 1788)
The
ssp. Erebia
pandrose orientalis Goltz, 1930 was described from the mountains of S.
Siberia, but this name was preoccupated by Erebia
epiphron orientalis Elwes, 1900. For this reason I have replaced this
name by yernikensis Korshunov, 1994,
nom. nov. It differs from the
butterflies of N. Europe by in general smaller size and more faded color of the
postdiscal field on the UPF.